By Sagar Shankaran, Founder of CallSphere
An in-depth look at Mixture of Experts (MoE) architecture, explaining how sparse activation enables trillion-parameter models to run efficiently and why every major lab has adopted it.
Key takeaways
The biggest LLMs of 2026 are not just larger -- they are architecturally different from their predecessors. Mixture of Experts (MoE) has become the dominant architecture pattern, powering models from Google (Gemini), Mistral (Mixtral), and reportedly OpenAI and Meta. Understanding MoE is essential for anyone working with or deploying large language models.
In a standard dense transformer, every token passes through every parameter in every layer. A 70B parameter model uses all 70B parameters for every single token. This is computationally expensive and scales poorly.
MoE changes this by replacing the feed-forward network (FFN) in each transformer layer with multiple smaller "expert" networks and a gating mechanism:
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Input Token -> Attention Layer -> Router/Gate -> Expert 1 (selected)
-> Expert 2 (selected)
-> Expert 3 (not selected)
-> Expert N (not selected)
-> Combine Expert Outputs -> Next Layer
The router (also called a gate) is a small neural network that decides which experts to activate for each token. Typically, only 2 out of 8 or 16 experts are activated per token.
The key insight is sparse activation. A model can have 400B total parameters but only activate 50B per forward pass. This gives you:
Mixtral 8x7B demonstrated this powerfully -- it has 46.7B total parameters but only 12.9B active per token, matching or exceeding Llama 2 70B performance at a fraction of the inference cost.
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The gating mechanism is the most critical component. Common approaches include:
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Load balancing is a real engineering challenge. If all tokens route to the same 2 experts, the other experts waste capacity. Training includes auxiliary load-balancing losses to encourage uniform expert utilization.
| Model | Total Params | Active Params | Experts | Architecture Notes |
|---|---|---|---|---|
| Gemini 2.0 | Undisclosed (rumored 1T+) | ~200B | MoE | Multi-modal, proprietary |
| Mixtral 8x22B | 176B | 44B | 8 | Open weights, Apache 2.0 |
| DeepSeek V3 | 671B | 37B | 256 | Fine-grained expert granularity |
| DBRX | 132B | 36B | 16 | Databricks, fine-grained MoE |
The trend is toward more experts with smaller individual capacity (DeepSeek's 256-expert approach) and shared expert layers that process every token alongside the routed experts. Research into dynamic expert creation and pruning could enable models that grow and specialize over time without full retraining.
Sources: Mixtral Technical Report | DeepSeek V3 Paper | Switch Transformers

Written by
Sagar Shankaran· Founder, CallSphere
LinkedInSagar Shankaran is the founder of CallSphere, where he builds production AI voice and chat agents deployed across healthcare, hospitality, real estate, and home services. He writes about agentic AI, LLM engineering, and shipping voice agents that handle real calls in production.
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